Revision of the genus Trilophus Andrewes from the Oriental region (Coleoptera, Carabidae)

Abstract

The species 444 Discussion 495 Phylogenetic aspects 495 Zoogeographie aspects 502 Alphabetical list of the species 506 Acknowledgements 506 References 535 OF THE GENUS TRILOPHUS 43 \ often material belonging to the genus Trilophus. In 1994, I was unable to define the species more precisely. So, at that time all material was provisionally placed under the taxon Trilophus interpunctatus (Putzeys), and the need for revision of the group noted (Balkenohl 1994). MaterialThanks to the kindness of many curators and private collectors it was possible to base this investigation on 944 Trilophus specimens out of about 80 populations. The material consists of the available type material as well as material of different expeditions carried out between the beginning of this century and 1996. Although collected at many and very different localities of the Oriental region from north-west Pakistan toBali, the study is incomplete due to the limited material and the spotty collecting. For some populations of a species only a few specimens were available and in few cases only one specimen was available at all.Old type material is distributed over various public collections. Most of these specimens could be located and in addition, material seen by Putzeys and Andrewes. As usually, old type material consisted of syntypes. For this material a lectotype and paralectotype(s) have been designated and redescriptions performed for all of these species.Most of the old type material bears several labels. All label information is listed for each of those specimens. Because I could not decide in some cases which of the different locality spelling is valid, all information given on labels was listed as it appears on the labels. If the genitalia of old type material were prepared this was performed specimen by specimen to avoid any possibility of confusion of labels. The original pins have been reused and if possible the old paper cards have been gluedadditionally to the pins. This was done to keep all information available for future investigation.

Trilophidius gemmatus sp. n., a new species from Bhutan, with an updated identification key to the Asian species (Coleoptera, Carabidae, Scaritinae)

“…Although revised, it was emphasized the genus might be much more widely distributed over the Oriental region including mountainous habitats. This assumption was based on the following knowledge: 1. the genus evolved over tropical Africa with much more species (15), 2. especially the terrestrial fauna of the Oriental Region is still relatively unexplored, and 3. the sistergroup of Trilophidius, Trilophus Andrewes, 1927 is known to be distributed over Asia with twenty-eight species (Balkenohl 1999). The present contribution re ports firstly evidence for the distinctly wider distribution.…”

Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini)

“…So, the question of the taxonomical position of Clypeuspinus gen. nov. among the Clivinini is challenging. In contrast to the Scaritinae, phylogenetic assessments are available only for some Clivinini groups [e.g., Schizogenius Putzeys, 1846 (Whitehead, 1972), Trilophus Andrewes, 1927(Balkenohl, 1999] but no comprehensive phylogenetic assessment for the subtribe is available at present. Therefore, relationships are described historically and until today on the basis of similarities observed when genera and species are compared, morphologically.…”