“…
Schwartz (2011) expanded the discussion and application of these terms to the
Orthotylini, discussing the various configurations of the
DES.
Schwartz (2011) also noted that several genera of the
Orthotylini have a single spiculum that he homologized with the
DES. Until further evidence among additional members of the
Zanchius group is available, we treated the single spiculum of
Carvalhomiris as the
DES, despite not completely agreeing with
Cassis (2008) that the base of the
DES has an expanded keel.…”