“…In the last decade, several systematic studies of the water lily family were conducted by combining morphological and molecular evidence. Based on molecular and morphological evidence, multiple studies support Brachyceras, Anecphya, Nymphaea, Hydrocallis, Lotos as a monophyletic subgenera, respectively, in agreement with our results (Borsch et al, 2007;LÖHNE et al, 2007;Borsch et al, 2008;Taylor, 2008;He et al, 2018). The combined morphological and molecular analyses conducted by Borsch et al (2008) suggest that the genus Nymphaea is paraphyletic, with the subgenus Nymphaea being sister to a clade comprising the other subgenus and the Euryale-Victoria clade.…”