“…This problem arises due to the fact that the processes that regulate stream communities vary across spatial and temporal scales, and range from local biotic interactions to large-scale biogeographic history (Tonn, 1990;Poff, 1997). The role of local, instream environmental variables (e.g., depth, substrate size, current velocity) in structuring stream communities has been well studied (Minshall, 1984;Degani et al, 1993;Voelz et al, 1994;Townsend et al, 1997;Fairchild & Holomuzki, 2002). In recent years, researchers also have begun focusing on how larger scale regional and landscape-scale factors (e.g., annual discharge patterns, watershed area, landuse) influence aquatic communities (Corkum, 1992;Richards et al, 1997;Sandin & Johnson, 2000;Parson et al, 2003;Weigel et al, 2003).…”