“…Alternatively, a completely different pattern, involving two lines of re-colonization, is possible. In addition to the one from the East, a second front reached Central and Northern Europe from either Iberia ("brown bear paradigm"; Hewitt, 1999) or refugia on the British Isles (there is growing evidence of unglaciated areas on the British Isles during the last ice age: e.g., Sinclair et al, 1998;Stewart & Lister, 2001;Hän-fling et al, 2002;Murton & Lautridou, 2003;Hoarau et al, 2007;Remerie et al, 2009). The location of the contact zone is similar to that hypothesized for the butterfly Maniola jurtina , which, however, expanded only from southern refugia (not from the British Isles).…”