“…Mucor circinelloides was the first species used for the industrial production of γ-linolenic acid (GLA)-rich oil, a precursor for several biologically active compounds ( Somashekar et al, 2003 ; Ratledge, 2013 ). Over the years, several other oleaginous filamentous fungi have been identified from different sources, including soil, herbaceous plants and animal feeds ( Liu et al, 2010 ; Dey et al, 2011 ; Banu et al, 2017 ; Muniraj et al, 2017 ; Tonato et al, 2018 ). Particular interest has been placed on Mucoromycetes (formerly Zygomycetes) and Ascomycota species, such as Aspergillus niger, Cunninghamella echinulata , Thermomyces lanuginosus (formerly Humicola lanuginosa ), Umbelopsis isabellina (formerly Mortierella isabellina ), and Umbelopsis vinacea (formerly Mortierella vinacea ), because of their high lipid accumulation ( Papanikolaou et al, 2004 ; Fakas et al, 2008 , 2009 ; Meng et al, 2009 ).…”