“…Biotic distributions instead have been used to infer that the ancestral Snake River flowed through southeast Oregon to the California Pacific coast, via either the Klamath (Taylor, 1960(Taylor, , 1985 or Sacramento (Miller, 1965) Rivers (also see Wheeler and Cook, 1954). Although this hypothesis is well entrenched in the literature (Christiansen and Yeats, 1992;Malde, 1965Malde, , 1991Smith et al, 2000), its supporting biogeographic evidence has never been rigorously evaluated.…”