“…However, several studies have documented that pollinator‐mediated selection on floral traits varies in space and time (Sletvold & Ågren, 2010; Chapurlat et al ., 2015; Wu & Li, 2017; Wu et al ., 2018), reflecting that plants live in complex and variable biotic and abiotic environments. The variation in strength and form of selection could be due not only to variation in the interaction intensity with pollinators (Sletvold & Ågren, 2014, 2016; Emel et al ., 2017; Trunschke et al ., 2017), but also to variation in interactive effects of other environmental factors, such as herbivory (Gómez, 2003; Ågren et al ., 2013; Sletvold et al ., 2015), reproductive assurance (Teixido & Aizen, 2019), co‐occurring species (Caruso, 2000; Eisen et al ., 2020) and soil resource availability (Sletvold et al ., 2017; Wu et al ., 2022). However, most studies focus on pollination in isolation, and the combined effects of several potential selective factors are poorly documented (reviewed in Sletvold, 2019).…”