“…A pale grey to light brown homogeneous mudstone with intercalated sandstone is the main deposit of the Duho 4 Formation (Hwang et al, 1995;Kim and Paik, 2013). The Duho Formation produces a variety of marine invertebrate and vertebrate fossils, including mollusks (Kim and Lee, 2011;Kong and Lee, 2012), fishes (Ko, 2016;Ko and Nam, 2016;Kim et al, 2018;Nam et al, 2021;Malyshkina et al, 2022), and whales (Lim, 2005;Lee et al, 2012). Such a diverse fossil record has produced equally diverse paleoenvironmental interpretations during the deposition of the Duho Formation.…”