“…Systematic investigations have shown the influence of genetic and metabolic factors in the construction of the layers. In fact, it is probable that both an endogenous rhythm and fluctuating environmental factors contribute, their respective importance being species-specific (May & Buttrose, 1959;Buttrose, 1962;Frey-Wyssling & Muhlethaler, 1965;Jenner, 1982). The morphogenesis of helicoidal walls provides an exceptional example, the importance of which is twofold: it is an oscillatory and subcellular example, and the movement of the constituent subunits mimics the movement of the hands of a clock, characteristics which may justify a comparison with horologic systems.…”