“…In plants, hybridization in particular—often accompanied by polyploidy, or whole‐genome duplication (WGD)—has been emphasized as an important evolutionary process (e.g., Stebbins, ; Soltis and Soltis, ) and a major source of observed cytonuclear discordance due to chloroplast capture (e.g., Rieseberg and Soltis, ; Soltis and Kuzoff, ; Linder and Rieseberg, ). While such instances of discordance are typically observed at shallow phylogenetic levels (e.g., Huang et al., ), recent studies have shown that ancient hybridization events can produce lasting signatures of cytonuclear discordance (Folk et al., ; García et al., ; Morales‐Briones et al., ), even among major lineages of angiosperms (e.g., members of the rosid COM clade; Sun et al., ). We might expect ILS, as well, to result in persistent cytonuclear discordance in some cases, but this possibility has been poorly explored across the phylogeny of green plants.…”