“…ultrametric trees), in which no terminal taxon can have such a disproportionate influence. Including fossils in time‐calibrated trees for ASR requires precise evaluation of their phylogenetic position and fossil record sampling bias (Bapst, 2014), as well as their effect on the underlying evolutionary model of the trait of interest (Cascini et al, 2019; King & Lee, 2015; Litsios & Salamin, 2012; Wilson et al, in press) – both of which have been the subject of considerable debate in the paleontological community (Bapst, 2014; Soul & Wright, 2021). While the inclusion of both fossil and extant species generally improves ASR accuracy – especially when using discrete traits, which are less sensitive to sampling bias and model misspecification (Puttick, 2016; Soul & Wright, 2021), the effect of such a high discrepancy in distance between nodes and terminal taxa on ASR with time‐calibrated trees remains to be assessed.…”