We study the dynamics of metallic vertical stripes in cuprates within the three-band Hubbard model based on a recently developed time-dependent Gutzwiller approximation. As doping increases the optical conductivity shows transfer of spectral weight from the charge transfer band towards i) an incoherent band centered at 1.3eV, ii) a Drude peak, mainly due to motion along the stripe, iii) a low-energy collective mode which softens with doping and merges with ii) at optimum doping in good agreement with experiment. The softening is related to the quasidegeneracy between Cu centered and O centered mean-field stripe solutions close to optimal doping. PACS numbers: 71.45. Lr, 71.10.Hf, 78.30.Er The doping-dependent evolution from insulating behavior to a strange metal in the superconducting cuprates emerges dramatically in the normal state optical conductivity. Slightly doped cuprates show a small or no Drude peak and doping-induced transfer of spectral weight from the charge-transfer (∼2eV) to a mid-IR (MIR) band at ∼0.5eV [1,2]. Upon further doping the system progressively metallizes as is evident from the prominent Drudelike peak that develops at zero energy. A remarkable effect of doping is that the MIR band strongly softens and merges with the Drude peak, resulting in a feature that cannot be fitted by a conventional Drude model. A variety of alternative theories [3,4] have been proposed in order to describe this feature. Clearly the identification of this low-energy MIR (LEMIR) band is of paramount importance to understand the physics of these materials. The low-doping behavior has been explained in terms of the random-phase-approximation (RPA) electronic excitations of single-hole Hartree-Fock states in CuO 2 layers [5], but the moderate doping behavior (the softening of the LEMIR band) could not be explained due to difficulties with the HF ground state.The softening of the LEMIR band in La 2−x Sr x CuO 4 (LSCO) is accompanied by the appearance of another (much less discussed) band at 1.3eV [1,2,6]. This high energy MIR (HEMIR) band is well pronounced in optical absorption through LSCO thin films[2], and electron energy loss spectroscopy [6] where it develops as a function of doping. Moreover LEMIR and HEMIR are also detected in photodoped experiments on LSCO [7]. The HEMIR has not been clearly resolved by reflectivity in YBa 2 Cu 3 O 6+δ (YBCO), but a strong broad feature at the right energy appears in photodoped transmission experiments [8]. As far as we know no microscopic explanation of the HEMIR exists so far.Another important aspect of layered cuprates that has emerged in the last years is the rearrangement of doped holes in antiferromagnetic (AF) domain walls[9, 10, 11, [11,12,13,14] (d is the distance between charged stripes in units of the lattice constant), chemical potential [18,19], and transport experiments [20,21] as a function of doping have been explained in a parameter free way [16].In this work we investigate the optical conductivity in terms of linear excitations around the metallic meanf...