Paraclupea seilacheri sp. nov., un arenque de doble armadura (Clupeomorpha, Ellimmichthyiformes) de las calizas albianas de la cantera Tlayúa, Puebla, México

Abstract

Paraclupea seilacheri sp. nov. is described in this paper. This ellimmichthyiform fish from the Albian deposits of the Tlayúa quarry, near Tepexi de Rodríguez, Puebla, central Mexico, shows the diagnostic characters of the superorder Clupeomorpha (the otophysic connection, commissure of the supratemporal sensory canal passing through parietals, and the abdominal scute series), order Ellimmichthyiformes (parietals in contact to each other along the midline of the skull; a beryciform foramen in the anterior ceratohyal; three epurals in the caudal skeleton; and a series of predorsal scutes), and family Paraclupeidae (predorsal scutes with increasing sizes in anterioposterior order and postpelvic abdominal scutes with a strong ventral spine). Additionally, the characters of this new Mexican fish are distinctive and support the creation of a new species of the genus Paraclupea, these include a diamond shaped body moderately high; 18 predorsal scutes strongly ornamented with radiating ridges; most posterior predorsal scutes are subrectangular, wider than long; the abdominal scutes are spatula-like; the dorsal process of posttemporal bone is hypertrophied; and the second uroneural is short and distally reaches the end of the first uroneural. The present finding introduces the second species of the genus Paraclupea; a significative increase in the geographical distribution of this genus in the Northern Hemisphere, from eastern China to the southern North America; and the third North American Cretaceous paraclupeid after Tycheroichthys dunveganensis and Scutatoclupea applegatei.

An Eocene paraclupeid fish (Teleostei, Ellimmichthyiformes) from Bolca, Italy: the youngest marine record of double‐armoured herrings

“…(), Alvarado‐Ortega & Ovalles‐Damián (), Khalloufi et al . (), Murray & Wilson (), Bannikov (), Figueiredo & Ribeiro (, ), and Alvarado‐Ortega & Melgarejo‐Damián ().…”

“…, Figueiredo & Ribeiro ), Iberia ( Ezkutuberezi carmenae ; Poyato‐Ariza et al . ) and China ( Paraclupea chetungensis ; Chang & Grande ), but some taxa are also known from shallow marine deposits of Brazil and Mexico ( Codoichthys carnavalii , Paraclupea seilacheri , Scutatoclupea applegatei ; see Alvarado‐Ortega & Ovalles‐Damián ; Figueiredo & Ribeiro ; Alvarado‐Ortega & Melgarejo‐Damián ). Vernygora et al .…”

“…(), Murray & Wilson (), Bannikov (), Vernygora & Murray (), Figueiredo & Ribeiro (, ), Murray et al . (), Alvarado‐Ortega & Melgarejo‐Damián () and Marramà & Carnevale (). Chronostratigraphic scheme adopted and modified from International Chronostratigraphic Chart v 2016/12.…”

“…This clupeomorph clade currently includes about 40 species arranged in 19 genera spanning from the Early Cretaceous to the middle Eocene (e.g. Grande ; Alvarado‐Ortega & Melgarejo‐Damián ; Figueiredo & Ribeiro ). However, the diversity of this extinct lineage can be easily increased with the revision of a number of species traditionally assigned to the genera ‘ Clupea ’ or ‘ Diplomystus ’ (Grande ).…”

On the Albian occurrence of Armigatus (Teleostei, Clupeomorpha) in America, a new species from the Tlayúa Lagerstätte, Mexico

“…Abbreviations in the predorsal scute formula: C, complete predorsal scute series; I, incomplete predorsal scute series; the number of predorsal scutes in each species is shown in front of the draws; draws show the change in the shape of the scutes along the predorsal observed in species; light arrows point to posterior scutes; dark arrows show the hypothetic intraspecific change of the predorsal scutes. resemble those of the complete predorsal scute series observed in other ellimmichthyiforms, such as Ellimma, Ellimmichthys, and Paraclupea (Chang and Grande, 1997;Chang and Maisey, 2003;Alvarado-Ortega and Melgarejo-Damián, 2017; among others). Armigatus namouraensis, A. dalmaticus, and probably A. oligodentatus have more predorsal scutes (10 to 19) that cover a larger portion of the predorsal edge of the trunk, lie over all the predorsal bones, and reach a position closer to the occiput (see Forey et al, 2003, figures 51.A, 54;Murray et al, 2016, figure 2; figure 1).…”