“…Tooth wear studies similarly suggested that horses consumed varying amounts of browse in northwestern Europe and eastern Beringia, depending on site and time (Rivals et al, , 2014Rivals and Lister, 2016;Semprebon et al, 2016), as well as inhabiting environments with varying levels of forest . Studies of modern Equus and Bison also indicate incorporation of some browse into diets (Craine et al, 2015b;Gębczyńska et al, 1991;Guthrie, 1990;Hansen, 1976;Kowalczyk et al, 2011;Marshal et al, 2012;Painter and Ripple, 2012;Peden et al, 1974;Schulz and Kaiser, 2013;van den Berg et al, 2015;Waggoner and Hinkes, 1986;Xu et al, 2012). Many Equus populations are grazers that consume minimal browse (e.g., Hansen, 1976), but browse is a dominant food in some populations of wild/feral Equus species either yearly or seasonally (Marshal et al, 2012;Schulz and Kaiser, 2013;Xu et al, 2012).…”