“…a natural park), while more complex locations, such as sparsely-located sites, have received much less attention. Although a few applications have focused on sites of the latter type, including forest areas (Czajkowski et al, 2017) and riparian zones (Holland and Johnston, 2017), to the authors' knowledge there is no study directly focusing on agricultural systems. If we consider substitutability and directional issues, it can be readily inferred that distance-decay models may perform poorly compared to area-and density-based models when applied to these sparsely-located ecosystems (Lizin et al, 2016;Budziński et al, 2018).…”