Let E be a real Banach space. We study the Ornstein-Uhlenbeck semigroup P = {P (t)} t≥0 associated with the Ornstein-Uhlenbeck operatorHere Q ∈ L (E * , E) is a positive symmetric operator and A is the generator of a C 0 -semigroup S = {S(t)} t≥0 on E. Under the assumption that P admits an invariant measure µ ∞ we prove that if S is eventually compact and the spectrum of its generator is nonempty, thenThis result is new even when E = R n . We also study the behaviour of P in the space BUC(E). We show that if A = 0 there exists t 0 > 0 such thatMoreover, under a nondegeneracy assumption or a strong Feller assumption, the following dichotomy holds: either P (t) − P (s) L (BUC(E)) = 2 for all t, s ≥ 0, t = s, or S is the direct sum of a nilpotent semigroup and a finite-dimensional periodic semigroup. Finally we investigate the spectrum of L in the spaces L 1 (E, µ ∞ ) and BUC(E).