“…The middle ear region of the mammalian skull has been a rich source of characters for phylogenetic work (Archer, 1976;Coombs and Coombs, 1977;Novacek, 1986;Hunt, 2001;Wible, 2003;Giannini et al, 2006;Mead and Fordyce, 2009;O'Leary, 2010;O'Leary et al, 2013;Orliac, 2013) due in large part to the complex and variable nature of the petrosal bone, which forms the "roof " of the middle ear (specifically the medial and superior borders), and the associated auditory bulla, which forms its "floor" (ventral and lateral borders). For the study of many mammal species, access to the petrosal bone in situ in the skull is direct because the "floor" of the middle ear, the auditory bulla, which covers the petrosal (variably composed of different bones, including petrosal, ectotympanic, and entotympanic, among others), is standardized treatment of the skulls of extant artiodactylans, describing the relationship of the petrosal to surrounding bones across the clade, currently exists.…”