“…Structural differences lead to a subdivision into branched GDGTs (brGDGTs) and isoprenoid GDGTs (isoGDGTs) (Schouten et al, 2013 ). The brGDGTs are known to originate from bacteria in terrestrial environments (soils, peats, rivers, and lakes) (Weijers et al, 2007 , 2010 ; Tierney et al, 2012 ; Dirghangi et al, 2013 ; DeJonge et al, 2014 ; Freymond et al, 2017 ; Huguet et al, 2017 ; Naafs et al, 2017 ) and the isoGDGTs are from archaea within aquatic (marine and lacustrine) and terrestrial systems (Pancost et al, 2011 ; Tierney et al, 2012 ; Dirghangi et al, 2013 ; Bischoff et al, 2014 ; Naeher et al, 2014 ; Schubotz et al, 2018 ; Bale et al, 2019 ; Evans et al, 2019 ). Both GDGT types occurred across the whole lithological profile with isoGDGT-0, isoGDGT-1, brGDGT-I, brGDGT-II, and brGDGT-III being the most prominent ( Figure 3D ).…”