“…The convergence of Physarum finding the shortest path has been proved by Bonifaci et al [32]. Based on its foragingbehavior, so far, the Physarum has been used to solve many problems, such as finding the shortest path in directed or undirected network [33,34,35,36], designing and simulating transport network [37,38,39,40], natural implementation of spatial logic [41,42], and computer music [43,44,45]. In addition, the Physarum model can also find the shortest path under uncertain environment [46,47] in the real application [48].…”