“…In addition to extensive heterogeneity in its natural cover, the Lerma-Chapala-Santiago watershed includes several major urban centers, including the cities of Guadalajara (1.7 million inhabitants), Toluca (~750,000 inhabitants), Queretaro (~730,000 inhabitants), Aguascalientes (~700,000 inhabitants), Morelia (~640,000 inhabitants), and Guanajuato (~480,000 inhabitants) (INEGI, 2005). With many regional urban centers within the watershed and Mexico City just beyond its limits, the expansion of the built environment exerts a strong and constant pressure on natural land covers [48,49] and water resources [35,50,51]. Processes of urbanization, extensive and intensive agriculture, timber harvesting, and mining fragment the landscape, creating a motley patchwork of human and natural land covers across most of the watershed [3,52,53].…”