“…Other authors have observed a diversity in arousal-related modulations both between (Shimaoka et al, 2018; Williamson et al, 2015) and within (Aydin et al, 2018; Liang et al, 2020; Molnár et al, 2021; Schröder et al, 2020) sensory systems. Therefore future work will need to establish if other thalamic nuclei, or neuronal sub-types within the dLGN (Kerschensteiner & Guido, 2017; Piscopo et al, 2013; Rosón et al, 2019), are similarly modulated by pupil size dynamics. Finally, although the pupil size signal is often lauded as an index for internal modulations that occur in the absence of other overt behaviors, considering pupil size in the context of other behaviors such as locomotion (Reimer et al, 2014; Vinck et al, 2015) or eye movements (Schwartz et al, 2020) promises to reveal a more diverse coupling to neural measures.…”