“…In Egypt, the majority of insect groups had their morphological characteristics evaluated and the consequences of different nesting spaces examined, but the Coleoptera, the group of insects with the greatest number of species, had neither of these studies (Riad, 2019;Riad and Mahmoud, 2020). Tenebrionidae is probably the biggest group of requested Coleoptera in the desert habitat, with over 20.000 known species and nearly 2307 genera of overall conveyance (Booth et al, 1990;Löbl and Smetana, 2008;Riad and Mahmoud, 2020). In spite of the fact that among all groups of beetles, Tenebrionidae (darkling beetles) ranks 7 th in terms of species diversity, due to the degree of the scarcity of knowledge regarding the phylogeny and systematics of the group, its monophyly has been addressed, as well as those of the subfamilies and tribes that belong to it (Condamine et al, 2011;Riad and Mahmoud, 2020).…”