Species' ranges are dynamic, flexible, and capable of change through range shifts, contractions, and expansions (Andrewartha & Birch, 1954;Sexton et al., 2009). Range shifts can occur as the result of stochastic processes, whereby random individuals at the edge of the species' range (hereafter: range edge) slowly expand the range over time through random movements (Skellam, 1951). Yet, range shifts