“…Studies based on morphology, osteology, ecology and, to a lesser extent, on genetics, have all made contributions to the present understanding of the Barbus genus Granado-Lorencio, 1988, 1997;CollaresPereira and Madeira, 1990;Torralva et al, 1997;Aparicio and Sostoa, 1998;Chenuil et al, 1998Chenuil et al, , 2000Tsigenopoulos et al, 1999, Callejas andOchando, 2000;Tsigenopoulos and Berrebi, 2000). However, basic questions on its evolutionary differentiation remain unsolved (see Tsigenopoulos and Berrebi, 2000 and references therein).…”