Cratons, the ancient nuclei of continents that have been stable for billions of years, are underlain by keels of lithosphere with strongly melt-depleted compositions 1,2 . These cratonic keels may have formed either from partial melting in a mantleplume environment 3,4 , or alternatively by melting at shallow depths in a subduction zone, during the successive accretion of slabs of oceanic lithosphere 5 . The stability of cratonic keels has been attributed to a pervasive state of near-neutral buoyancy-isopycnicity-created by offsetting thermal and compositional effects on density 6 . However, it is unclear how an isopycnic state can be sustained over geological time 2 . Here we simulate the evolution of a simplified southern African cratonic keel, initiated in either a hot-plume or a cold-slab environment, over 3 billion years, using a numerical model that incorporates secular cooling of the mantle, coupled with gradual loss of radiogenic heating in the lithosphere. We find that the simulation that starts from a cold-slab environment best explains the subsidence history of the southern African craton 7 . However, irrespective of how the cratonic keel formed, we find that the isopycnic state is inherently ephemeral: a cratonic keel that is approximately isopycnic under present conditions was more, or less, buoyant in the geologic past.The lithosphere, Earth's relatively rigid outer shell, moves coherently with plate motions above the underlying, more easily deformed asthenosphere. The lithosphere-asthenosphere boundary (LAB) thus forms a mechanical detachment that accommodates relative motion between the plate and underlying mantle; as such, it represents the most extensive plate boundary on Earth 1 . Yet, owing to the scarcity of representative geological samples 8 , even fundamental physical parameters remain uncertain, particularly beneath Archaean cratons 9 . The mantle underlying these regions may be compositionally stratified 10 , and possesses a distinctive, dehydrated and strongly melt-depleted composition that differs fundamentally from modern analogues 3 . This mantle region, sometimes denoted as tectosphere 6 , is stabilized against convection under present mantle conditions owing to its intrinsic buoyancy and relatively high viscosity 2 . The buoyancy and rheology of cratons under thermal conditions in the geologic past are unknown, however, particularly during the early stages of lithospheric evolution.The formation of cratonic lithospheric mantle (CLM) is contentious and various models have been proposed, of which two may be considered as thermal endmembers. One model invokes formation of CLM within a hot Archaean plume environment, where CLM peridotites represent residues and/or cumulates from high-degree partial melting at significant depth 3,4 . A second model postulates formation of CLM by melting at shallower depths, associated with underthrusting and imbrication of subducted oceanic lithosphere 5 . Both of these models attribute longevity of CLM to intrinsic buoyancy arising from a strongly melt...