Miamia maimai n. sp., a new Pennsylvanian stem-orthopteran insect, and a case study on the application of cladotypic nomenclature

A new, rare and small “lobeattid” species (Insecta: Archaeorthoptera) found at Xiaheyan (Pennsylvanian; Ningxia, China)

“…Non-panorthopteran Archaeorthoptera include "lobeattid" insects, the order Cnemidolestodea Handlirsch, 1937 (which is probably derived from a subset of lobeattid insects) and a number of unplaced species. Béthoux, 2008;Béthoux et al, 2012b; Jarzembowski, 2010) a comparatively broad ScP-R/RA area. Another peculiar trait shared with the corresponding species (but not unique to Miamia spp.)…”

“…Many species could be documented on the basis of large to very large samples, in particular among the stem lineages of Orthoptera (crickets, katydids and grasshoppers), which dominate the various taphocoenoses from which insect remains were recovered (Trümper et al, 2020). For example, species such as Longzhua loculata Ren, 2011, Miamia maimai Béthoux, Gu, Yue andRen, 2012b and Protomiamia yangi Du, Béthoux, Gu and Ren, 2017 were described based on several tens of specimens each. However, the undertaken collecting effort also allowed sampling of rare species of stem Orthoptera, including Heterologus duyiwuer Béthoux, Gu and Ren, 2012a and Sinogerarus pectinatus Gu, Béthoux and Ren, 2017, each known from a single, isolated forewing.…”

The earliest and most oriental Calvertiellidae unearthed (Palaeodictyoptera; Late Carboniferous; China)

“…Instead we suggest that by the early Late Carboniferous many insect lineages of comparatively moderate importance (i.e., considered of familial level or below) had already undergone dispersion at a large scale. Not only the palaeodictyopteran families Spilapteridae and Calvertiellidae show a wide distribution, but also particular genera of the Megasecoptera (Pecharová et al 2015), stem-Orthoptera, such as Miamia spp., recovered from at least three Late Carboniferous localities (Béthoux 2008;Béthoux & Jarzembowski 2010;Béthoux et al 2012), and the Cnemidolestodea (Béthoux 2005;Aristov 2012;Gu et al 2014, among others). The Odonatoptera are no exception, with genera shared among many localities worldwide (Li et al 2013a), some ranging from the early Late Carboniferous to the Permian.…”

A new, rare and distinctive species of Panorthoptera (Insecta, Archaeorthoptera) from the Upper Carboniferous of Xiaheyan (Ningxia, China)

“…This exceptional material allowed solid inferences on wing venation intra-specific variability in various insect lineages (e.g. Béthoux et al, 2011Béthoux et al, , 2012bCui et al, 2011;Gu et al, 2011;Li et al, 2013;Pecharová et al, 2015). Moreover, both males and females were identified for two species (Du et al, 2017;Pecharová et al, 2015), one of them, a relative of grasshoppers, crickets and katydids (Orthoptera), displaying a significant femalebiased sexual size dimorphism.…”