“…Both Paracaryum and Solenanthus are, in turn, closely related to Cynoglossum L., a broad genus, the limits of which have long been a source of controversy (Mill 1979). The lack of a satisfactory correlation between floral and fruit characters in this group of Cynoglosseae, which is possibly due to parallel evolution, led Greuter (1981) to merge Rindera, Solenanthus and Paracaryum with Cynoglossum, in the wait for additional evidence to elucidate the phylogenetic relationships in the group. This treatment, however, is in contrast with that of most students of Boraginaceae (e.g., Kusnetsov 1910, Brand 1921, Johnston 1924, Popov 1950, 1953, Mill 1979, who kept the above genera separate (except Johnston who placed Solenanthus under Cynoglossum).…”