“…High PC2 values reflect greater variation in seasonal temperatures and lower total annual precipitation, which typify areas of especially high C. solstitialis density in California (Dlugosch, Cang, et al, ). Previous studies in this system have proposed that C. solstitialis success stems from a lack of effective competitors in more drought prone habitats (Dlugosch, Cang, et al, ), due in part to the extensive conversion of these habitats to rangeland (Menke, ; Stromberg & Griffin, ). Other studies within the California invasion, however, have found that water availability (both naturally occurring and experimentally manipulated) is strongly and positively correlated with C. solstitialis density and fecundity (Enloe, DiTomaso, Orloff, & Drake, ; Eskelinen & Harrison, ; Hulvey & Zavaleta, ; Morghan & Rice, ), suggesting that fitness should be highest in wetter areas.…”