<p><strong>First country record of </strong><strong><em>Pseudoathyreus orientalis </em></strong><strong>(Coleoptera: Bolboceratidae: </strong><strong>Athyreinae) from Pakistan</strong></p>

Abstract

Bolboceratidae is a large family of about 600 species (e.g., Schoolmeesters 2019), in the superfamily Scarabaeoidea. Currently, it includes two subfamilies, Athyreinae and Bolboceratinae (e.g., Scholtz &amp; Grebennikov 2016; Nikolajev et al. 2016). The Athyreinae includes five genera and about 130 species from the Afrotropical, Neotropical, Oriental and Palearctic regions (e.g., Schoolmeesters 2019). The genus Pseudoathyreus Howden &amp; Martínez, 1963 contains nine species occurring in Africa and Asia (e.g., Carpaneto et al. 2000; Schoolmeesters 2019). Little is known about the biology of Athyreinae (Scholtz &amp; Grebennikov 2016). Immature stages of this group remain unknown (Grebennikov &amp; Scholtz 2004).