2. Study Site LPAH is a small (1.2 km 2), shallow (avg. depth <1 m, max. depth 1.5 m), alkaline lake in The PNWR, a 17.7 km long, 21.8 km 2 area of marshes, impoundments, and shallow lakes that lies at the southern end of the Pahranagat Valley in southern Nevada (Fig. 1A and B). LPAH is spring-fed, receiving water from a number of springs that discharge north of the lake, then owing south through the Pahranagat Valley (Fig. 1A). The deep regional carbonate aquifer extends north to Great Basin National Park, east into Utah, and south to Death Valley and Lake Mead (Eakin 1963). U.S. Geological Survey stream-gages for the two highest discharge springs, Crystal, and Ash discharged at 340 l s-1 and 510 l s-1 between 2004-09 with discharge at a relatively constant rate throughout the year (Paces and Wurster, 2014). The Pahranagat Valley is in a region with some of the highest evaporation rates in the U.S. Wurster (2010) calculated that open water evaporation approaches 1676 mm yr-1 , greater than 10 times the average annual precipitation.