“…This has been studied in wheat since 1937 (Bayles, Taylor & Bartel, 1937), and in excised wheat leaves since 1969 (Salim, Todd & Stutte, 1969) as a technique for measuring drought avoidance in cereal seedlings. Since then, excised-leaf water loss (ELWL) in wheat has been used extensively as a selection tool indicating drought tolerance (e.g., Dedio, 1975; Kirkham et al, 1980; Clarke & McCaig, 1982b; Winter, Music & Porter, 1988; Yang, Jana & Clarke, 1991; Dhanda & Sethi, 1998; Golestani Araghi & Assad, 1998; Chandra & Islam, 2003; David, 2010). Thus, rate of water loss from excised leaves has been negatively associated with grain yield under drought in wheat in many reports (e.g., Clarke & McCaig, 1982a; Clarke & Townley-Smith, 1986; Clarke et al, 1989; Haley, Quick & Morgan, 1993; Chandra & Islam, 2003).…”