“…Low tissue mass density in leaves is associated with a high specific leaf area, large total leaf area and fast growth, on the one hand (Garnier, 1992 ;Ryser & Lambers, 1995 ;Meerts & Garnier, 1996 ;Ryser & Aeschlimann, 1999), and with short life span and high resource losses on the other (Schla$ pfer & Ryser, 1996 ;Ryser & Urbas, 2000). Leaf toughness and high tensile strength are associated with slow growth and also with a high degree of mechanical stability of the leaves, which tend to be well protected against environmental hazards such as herbivory (Coley, 1983 ;Reich et al, 1991 ;Cornelissen et al, 1999). Hence, tissue mass density is an effective means of predicting the performance of plants along gradients of resource availability (Wilson et al, 1999) ; a low tissue mass density is characteristic of plants of productive habitats, a high tissue mass density is typical in plants of unproductive environments Studies on the ecological significance of tissue structure have mostly been restricted to aboveground organs, especially leaves.…”