“…Pollen, plant macrofossil and ancient DNA studies suggest that the mammoth steppe was a highly productive herb-steppe-tundra environment, dominated by grasses, sedges, forbs and herbaceous species, with some shrubs such as willow also present. There is little evidence for trees during stadials; however the region contained forests during interstadial and interglacial times (Binney et al, 2017;Burns, 1991;; Guthrie, 1990Castaños et al, 2014;Huntley et al, 2013;Mandryk, 1996;Muhs et al, 6 2001; Schweger et al, 2011;Willerslev et al, 2014;Zimov et al, 2012). Plant species were unevenly distributed across the mammoth steppe due to local effects such as topography, drainage and latitude, and thus formed a "vegetational mosaic" .…”