“…For ten other isozyme loci examined, intraspecific variation in banding patterns in the allotetraploid species largely reflected the interspecific variation present in their putative diploid progenitors, and was offered as evidence for multiple, independent origins of the allotetraploid species (see section Multiple Origins of Hybrid Fern Taxa). Isozyme electrophoresis was subsequently used to identify the progenitors of allopolyploid species and hybrids in numerous temperate and tropical genera, including Dryopteris (Darnaedi et al, ; Werth, ; Jiménez et al, ; Stein et al, ), Cystopteris (Haufler et al, ; Haufler & Windham, ), Polypodium (Bryan & Soltis, ; Haufler & Windham, ; Haufler et al, ), Adiantum (Paris & Windham, ), Pellaea (Gastony, ), Hemionitis (Ranker et al, ), Polystichum (Soltis et al, , ; Barrington, ), Gymnocarpium (Pryer & Haufler, ), Botrychium (Hauk & Haufler, ), and other groups of Asplenium (Herrero et al, ; Van den heede et al, , ).…”