“…Analysis of each tool-use event provided a record of the grips used to manipulate the stone tool, how long individual grips were used, and a total overall record of how long each tool-use event lasted. Given that individual tools were used for varying lengths of time due to external variables, such as biometric differences across participants or differing tool-use contexts (Key andLycett, 2011, 2017c, in press;Merritt, 2012Merritt, , 2016), it was not always possible to directly compare time records across different tool types and forms. Therefore, we calculated an additional variable, 'Percentage of Use' (PoU), which was the percentage of time that a specific hand grip was used relative to the overall time record for which that tool was used.…”