Boessenecker et al., 2019). Considering the significant role that modern sharks play in regulating prey communities and ecosystem structure (Ferretti et al., 2010), its extinction likely had a profound impact on the functioning of the ecosystem and the dynamics of the trophic web within ancient marine communities (see Pimiento & Clements, 2014; Cooper et al., 2022). Conventionally, it has been hypothesised that † O. megalodon was constrained to warm temperatures,