“…In animal ecology, however, the process must also ensure that the chosen metrics capture all dimensions of spatial pattern relevant to an animal. For exploratory studies of the influence of mosaic pattern on animals, a suite of metrics is typically used (26 metrics by McGarigal and McComb (1995) for bird assemblages; 17 metrics by McAlpine et al (1999) for kangaroos and eight metrics by Hansen et al (2001) for caribou). For marine systems, however, this type of approach remains largely theoretical (Robbins and Bell, 1994), with few attempts to quantify spatial pattern (Garrabou et al, 1998 for rocky shores; Robbins and Bell, 2000 for seagrasses; Teixidó et al, 2002, for Antarctic benthic communities).…”