“…By studying the foraging ecology of three European large herbivore species over millennial timescales during the Holocene we aimed to: (a) identify habitat preferences and diets before and after the Neolithic environmental transformations; (b) analyse the response of the largest remaining Pleistocene megafauna in Europe to the major environmental changes of the Holocene (from increasing tree cover during the Early and pre-Neolithic Holocene to increasing habitat openness during the Neolithic and subsequent time periods); and (c) identify factors influencing their patterns of habitat use and diet. We hypothesized that forest expansion in the early Holocene forced large herbivores, especially those primarily adapted to grazing (aurochs and European bison) into forests as refugial habitats, to shift their pattern of habi- Daugnora, Jankauskas, & Ogrinc, 2009;Bocherens et al, 2015;Fornander, Eriksson, & Liden, 2008;Gravlund et al, 2012;Jessen et al, 2015;Lidén, Eriksson, Nordqvist, Götherström, & Bendixen, 2004;Linderholm, Fornander, Eriksson, Mörth, & Lidén, 2014;Noe-Nygaard et al, 2005). In total, 364 large herbivore bone samples and literature records were used in the study (Figure 1).…”