“…Major factors determining ichnological features (i.e., abundance, composition and diversity of trace fossil assemblages) in a deep-sea setting are food availability, bottom and pore-water oxygenation, substrate consistency, and rate of sedimentation (Wetzel, 1991;Uchman et al, 2008Uchman et al, , 2013aRodríguez-Tovar et al, 2009a, b;Rodríguez-Tovar and Uchman, 2010;Uchman and Wetzel, 2011;Wetzel and Uchman, 2012;Rodríguez-Tovar and Reolid, 2013;Rodríguez-Tovar and Dorador, 2014 , 1984;Ekdale, 1992;Schlirf, 2000), and Planolites, an actively filled burrow, is interpreted as a pascichnion in shallow tiers (see Pemberton and Frey, 1982;Keighley and Pickerill, 1995 the latter is interpreted as a shallow tier, pascichnia structure, in deep-marine, low energetic, oxygenated, environments (Uchman, 1995;Mángano et al, 2002;Wetzel, 2002;Löwemark et al, 2012), associated with increase food flux, feeding on microbes that occurs in high concentrations (Wetzel, 2002;Löwemark et al, 2012). tracemaker (see Löwemark and Werner, 2001;Bromley andHanken, 2003, Löwemark andSchäfer, 2003;Löwemark et al, 2004;Löwemark, 2015).…”