“…Concentrations of K + , Ca 2+ , and Mg 2+ in the throughfall were high compared to those in the BD with NTDs of 0.97, 0.13, and 0.13 kg ha -1 mo -1 (3-year average), respectively. This result is consistent with those from other studies that have shown that, in various forest types, a large proportion of the ions in the throughfall originated from canopy leaching (Cronan et al, 1983;Parker, 1983, Okochi et al, 1995Houle et al, 1999;Muramoto et al, 2007). As shown in Figure 4, NTDs of most ions except Na + increased as the drenched duration of the canopy increased.…”