“…Airborne pollen concentrations are measured to be anywhere between 10 and 1000 grains m −3 (Després et al, 2012) but can reach between 10 4 and 10 6 grains m −3 , particularly in the Northern Hemisphere spring months of April and May when many deciduous trees display peak pollen emissions (Steiner et al, 2015;Gregory, 1978). Pollen grains have been shown to reach altitudes of 3 km and above in the atmosphere (Diehl et al, 2001) and travel horizontal distances of up to ∼ 10 3 km (Sofiev et al, 2006;Hjelmroos, 1992;Sack, 1949). Individual small pollen grains can reside in the atmosphere for a few (generally 2-4) days (Sofiev et al, 2006), although pollen seasons can persist with high average concentrations (∼ 10 4 grains m −3 ) over many months, fluctuating with locality, season, time of day, and meteorology (Gregory, 1978).…”