“…Unfortunately, the poor situation in SE Spain could become the common situation in Europe and North Africa, where many Z. lotus populations occur in coastal areas (Figure 1; Mücher, Hennekens, Bunce, Schaminée, & Schaepman, 2009) and are threatened by urban and agricultural sprawl, climate warming, aridification, and seawater intrusion (Ibáñez, Pérez-Gómez, Oyonarte, & Brevik, 2015). The compulsory registration and protection (Articles 1 and 6 and Annex 4.5 of the WFD) of GDEs and the sexennial report on the conservation status of habitats (Article 17 of Habitats Directive) should urgently include an assessment of the dependence of Habitat 5220* on groundwater.Indeed, identifying GDEs is an increasingly demanded step in Europe(Boulton, 2005;Kløve et al, 2011) for the protection and management of GDEs under the WFD (Articles 1 and 6) and a first step for the assessment of the hydrological ecosystem services(Carvalho-Santos et al, 2013) provided by GDEs. This identification becomes a challenge when groundwater is not observed on the surface at any time of the year (U.K. Technical Advisory Group, 2012), and our approach can assist with this purpose.…”