In the course of the evolution of the genus Homo, the most profound developments in life history parameters seem to have occurred in the Lower Pleistocene. Yet Acheulian industries are widely seen as having remained essentially unchanged for some 1.3 million years or more. In reality, however, although the Acheulian did not develop in a cumulative or directional manner over its long history, it nevertheless displayed considerable levels of typological and technological diversity and variability at continental, regional, and local levels. It is at the local level that this variability is at its greatest, with prepared core technologies regarded as characteristic of the succeeding Middle Paleolithic and Middle Stone Age appearing sporadically and ephemerally in the Acheulian. It is our contention that this pattern of local, short term variability combined with global long term stasis cannot be accounted for by models asserting that the hominin makers of the Acheulian lacked the cognitive capabilities of their evolutionary successors. Instead, we argue that Acheulian hominins were cognitively capable of innovative technical behavior and often displayed it; but that, despite structural life history parameters that approached those of living Homo sapiens, relatively short childhood, juvenile, and adolescence phases, combined with small local group size, constrained the social and developmental scope for innovation. Furthermore, we argue that metapopulation-level social, demographic, and ecological dynamics in the Acheulian, relating to group size, foraging ranges, and levels of individual migration, served to limit the lifespan of local groups and thereby reduced the likelihood of innovative behaviors disseminating through social networks and becoming fixed in cultural repertoires before the originator population became extinct. We explore the idea that the transition from the Acheulian to the ensuing Middle Paleolithic and Middle Stone Age was therefore driven not by evolutionary developments in hominin cognitive capacities, but by changes in life history and metapopulation factors.
INTRODUCTIONI n the course of the evolution of the genus Homo, the most profound developments in life history parameters seem to have occurred in the Lower Pleistocene (papers in Thompson et al. 2003). Yet the Acheulian industries that dominate the archaeological record of Africa and large parts of Eurasia from around 1.6 million years ago appear to have remained largely unchanged typologically and technologically for some 1.3 million years or more, maintaining through that timespan and geographic range the essential signature of large cutting tools (LCTs)/handaxes combined in varying proportions with a simple core-and-flake component. So striking is this apparent technological stasis that the Acheulian has often been described as 'monotonous,' 'stagnant,' and as a 'long oscillation' (Isaac 1972(Isaac , 1976 with no directional trend (Leakey 1975). Yet, on closer examination, the archaeology of the Acheulian furnishes evidence fo...