“…However, such an offset is not unprecedented. In the Cenozoic, TEX 86 generally agrees well with the trends and absolute temperatures reconstructed from alkenones (e.g., U K′ 37 ), foraminiferal δ 18 O and Mg/Ca ratios, and clumped isotopes (Δ 47 ) of carbonates, though commonly showing a slight offset to warmer SSTs (Bijl et al, 2009(Bijl et al, , 2010Castañeda et al, 2010;Douglas et al, 2014;Hollis et al, 2009Hollis et al, , 2012Huguet et al, 2006;Keating-Bitonti et al, 2011;Lopes dos Santos et al, 2013;Menzel et al, 2006;Meyer et al, 2018;Naafs et al, 2018;Pearson et al, 2007;Richey et al, 2011;Shintani et al, 2011;Woelders et al, 2018;Zachos et al, 2006). The causality of these higher temperatures remains unresolved but has been attributed by many authors to a summer bias of the Thaumarchaeota or a winter bias of the other proxies.…”