“…Ecosystem-atmosphere CO 2 exchange in western forests is controlled in large part by the annual delivery of water from the melting winter snowpack (Hu, Moore, Burns, & Monson, 2010;Monson et al, 2005;Winchell, Barnard, Monson, Burns, & Molotch, 2016), while summer rains can compensate for snowpack limitations (Berkelhammer, Stefanescu, Joiner, & Anderson, 2017;West, Hultine, Burtch, & Ehleringer, 2007). Summer rain in the southwestern United States is delivered by the North American Monsoon (NAM), a climate system that has been active during the Holocene (Betancourt, 1990;Bhattacharya, Tierney, & DiNezio, 2017). The utilization of soil water from different seasonal precipitation sources, and the photosynthetic efficiencies that are realized by interannual variation in the amounts of winter and summer precipitation, can be inferred through studies of the carbon and oxygen stable-isotope ratios in the α-cellulose of tree rings (Barbour, Walcroft, & Farquhar, 2002;Craig, 1954;Francey & Farquhar, 1982;Leavitt & Long, 1986;Libby & Pandolfi, 1974).…”