“…The chemistry of Al-humus complexes also plays a major role in soil pH buffering and in the regulation of Al activity in soil solution (Mulder and Stein, 1994;Takahashi et al, 1995). The Al bound to organic complexes is of low toxicity (Adams et al, 2000;Matús et al, 2006), and this may explain the lack of negative effects of Al on the growth of tree species on acid soils.…”