“…A considerable amount of research has focused on nutrient limitation in young and mature lodgepole pine trees, often within a silvicultural context, with different results reported in different settings (e.g., Yang, 1998 and references cited therein;Brockley, 2000Brockley, , 2003Startsev et al, 2007). Although N has often been found to be limiting, growth also may be limited primarily by other nutrients such as sulfur or boron (e.g., Brockley, 2000Brockley, , 2003Kishchuk et al, 2002;Blevins et al, 2005;Sanborn et al, 2005).…”