“…The first group, which Peppe et al (2007) named the "brevis-type," was similar to the generic type, whereas a second group, which they named the "tenuinervis-type," had a thinner texture, was often entire or nearly entire with irregularly spaced lobes or pinnules, and lacked the transverse corrugations and the circinate vernation of the brevis-type. However, despite their disparate morphologies, both groups share cuticular characteristics, both can have resin bodies between their veins, and both have their laminar tissue attached to, and entirely covering the midvein-a character unique among cycads (Nathorst 1909;Seward 1917;Harris 1964;Watson and Cusack 2005).…”