“…Trace fossils are typically the only means of evaluating biodiversity in ancient terrestrial deposits, particularly in paleosols, where the preservation potential of body fossils is typically low but organism abundance can be high (e.g., Retallack, 1984;Genise et al, 2000Genise et al, , 2002Hasiotis et al, 2003;Hasiotis, 2004Hasiotis, , 2007Hasiotis, 2007a, 2008;Hembree and Nadon, 2011). Trace fossils are generally easily preserved even in environments where body fossils are rare; therefore, the use of both body and trace fossils are necessary to provide the best paleoecological interpretations (e.g., H€ antzschel, 1975;Osgood, 1975;Frey, 1978;Bromley, 1996;Pemberton et al, 2001;Hasiotis, 2007).…”